Table of Contents
- Definition of a Lipid
- Dietary Lipid Occurs in Three Physical Forms: Fat Droplets, Membranes,
and Skin Lipids
- Fat (Mostly Triglyceride) is a Major Source of Calories in a Typical
U.S. Diet
- Fat Droplets
- Average Daily Input of Lipids to the Gastrointestinal Tract of Western
Man
- Bile is a Source of Endogenous Lipid and Detergent (Bile Salt)
- Fat Absorption is 98% Efficient Over a Wide Range
- The Composition of Fecal Fat in Normal U.S. Adults
- Neurohumoral Control of Digestive Secretions
- Release and Action of Cholecystokinin
- Steps in Troglyceride Absorption
- Triglyceride Digestion, Absorption, Resynthesis, and Secretion Occur
Simultaneously
- Key Physicochemical Events in Lipid Digestion
- Multiple Lipases Assure Efficient Intraluminal Hydrolysis of Dietary
Lipid
- Lipolytic Enzymes Act in Small Intestine at Micellar Bile Salt Concentrations
- Fat Absorption Persists in the Absence of Billary or Pancreatic Secretions
- The Relative Importance of Different Lipases for Triglyceride Digestion
Chnages With Age
- The Relative Importance of Acid Lipases in Fat Digestion in Cases of
Pancreatic Insufficiency
- Pancreatic Carboxyl Esterase Hydrolyzes a Variety of Lipid Esters
- Pancreatic Phopholipase A2 Hydrolyzes Membranes Phospholipids
- Differing Positional Specificity of Triglyceride Lipases
- Perferred Physical Forms of Substrates for Lipolytic Enzymes
- pH Optima of Gastrointestinal Lipases
- 10-30% Fat Digestion Occurs With Action of Acid Lipases in Stomach
- Fatty Acids Produced During Gastric Lipolysis Help Emulsify Fat in the
Stomach
- Emulsification of Fat in the Stomach: Gastric Content Being Propelled
Retrograde
- Fatty Acids Produced in Stomach Augment Duodenal Lipolysis by Multiple
Mechanisms
- Properties of Pancreatic Lipase
- Effect of Bile Salts and Colipase on Pancreatic Lipase Activity
- Bile Salts Disperse the Products of Fat Hydrolysis
- Visible Product Ohases Can Be Seen During Fat Digestion on a Microscope
Slide
- Fat Droplets Undergoing Digestion on a Microscope Slide
- The First Seconds of Lipolysis
- Bile Salts Disperse the Lipolytic Product Phases
- Bile Salts Solubilize Products of Lioplysis in Mixed Micelles
- Centrifugation of Intestinal Content Partially Separates the Phases
Present During Fat Digestion
- Monomeric and Micellar Lipids and Bile Salts are in Dynamic Equilibrium
- Bile Salts Micelles Greatly Increase the Capacity of Water to Carry
Fatty Acids
- Micellar Solubility of Lipolytic Products of Long Chain Triglyceride
in Bile Salt Solution
- Intestinal Mixed Micelle Contains Bile Salts and Two Classes of Lipid
Solute
- Micellar Solubilization of Secondary Solutes is Influenced by Multiple
Factors
- Solubilized Dietary Fat Can Greatly Enhance Absorption of Otherwise
Poorly
Absorbed Secondary Micellar Solutes
- A Fluorescent Carcinogen Dissolved in Fat Moves into Product Phases
During Lypolysis
- Unabsorbed Oils Trap Secondary Solutes and Reduce Their Absorption
- Plant Sterols Competitively Inhibit Cholesterol Solubility in Mixed
Micelles
- Lipid Absorption is Limited by Slow Diffusion of Micelles and Individual
Molecules Through the Unstirred Layer
- Unstirred Layer Thickness is Decreased by an Increase of Flow
- Effect of Micellar Solubilization of Fatty Acid on Diffusive Flux Through
Unstirred Layer
- Possible Mechanisms of Membrane Transport of Fatty Acids
- Rate and Site of Fat Absorption is Modulated by Presence of Micelles
- Differing Sites and Mechanisms of Fat and Bile Salt Absorption
- The Luminal Face of the Intestinal Microvillus Membrane is Packed with
Glycolipids and Cholesterol and Thereby Resistant to Digestion by Phopholipase
A2
- Structure of the Microvillus Cytoskeleton Through Which Lipolytic Products
Must Diffuse
- Luminal Contents During Fat Digestion
- Fatty Acid Binding Proteins May Assist in Intracellular Transport of
Absorbed Fatty Acids
- High Intracellular Concentrations of Toxic Fatty Acids in Enterocyte
Are Avoided By:
- Chylomicron Formation and Secretion
- Two Pathways of Triglyceride Synthesis in Intestine Insure Reesterfication
of all Dietary Fatty Acid
- Structure and Composition of Chylomicron
- Chylomicrons Are Too Largeto Enet the Pores in Capillaries and Therefore
Leave the Intestine Via the Lymphatic System
- Chylomicron Bypass Liver and Enter Systematic Circulation Via Thoracic
Duct
- Digestion and Absorption of Medium Chain Triglyceride (MCT)
- Salad Oils May Contain Small Amounts pf Fat Soluble Carcinogens
- Fate of Fat Soluble Drugs and Xenobiotics is Influenced bby Biotransformation
in Enterocytes
- Fate of Fat Soluble Drugs and Xenobiotics is Influenced bby Biotransformation
in Enterocytes
- Newly Resynthesized Fat in the Intestine Conatins Fluorescent Molecules
Originally Dissolved in the Dietary Fat
- Function of Intraluminal Bile Salts
- Physicochemical Properties of Conjugated Bile Salts Promote High Intraluminal
Concentrations During Digestion
- Only Terminal Ileum Actively Absorbs Bile Salts
- Weakly Polar Molecules Are Passively Absorbed
- Important Differences Between Conjugated and Unconjugated Bile Salts
- Bile Salts Kinetics in Man
- High Aqueous Concentrations of Bile Salts and Fatty Acids in the Colon
Are Toxic
- Multiple Factors Keep the Aqueous Concentration of Colonic Bile Salts
Low
- Colonic pH Influences Physical State and Cathartic Activity Malabsorbed
Bile Salts
- Bile Salt Malabsorption Causes Decreased Billary Secretion When Maximal
Synthesis Cannot Compensate for Increased Loss
- Factors Causing Decreased Luminal Concentration of Mixed Micelles
- Different Intraluminal Defects Causr Fat Malabsorption
- Mechanisms of Fat Malabsorption
- Different Mucosal Defects Cause Fat Malabsorption
- Digestion of Triglycerides is Influenced by the Length of the Fatty
Acid Chain
- Mucosal Processing and Body Metabolism of Fatty Acids Depends Upon Their
Chain Length
- In Lipase Deficiency, There Are Few Lipolytic Products to be Solubilized
by Bile Salts
- In Bile Salt Deficiency, Lipolytic Products Are Formed But Poorly Dispersed
- Fatty Acids Induce Net Fluid Secretion (Diarrhea)
- Non-Absorbed Dietary Fatty Acids Can Induce Colonic Fluid Secretion
- Causes of Impaired Triglyceride Hydrolysis
- There is Normally a Large Excess of Lipase Secreted for Fat Digestion
- Bile Salt Malabsorption Causes Progressive Bile Salt Deficiency Throughout
the Day
- Consequences of Lipase Deficiency
- Consequences of Bile Salt Deficiency
- Steatorrhea After Ileal Resection is Influenced by Digestive and Absorptive
Factors
- Solubilization of Lipolytic Products is Similar in Vitro and in Patients
- Dietary Oxalate Absorption is Enhanced and Prolonged in Patients with
Steatorrhea Due to Bile Salt Deficiency
- Apparent Mechanism of Oxalate Hyperabsorption in Steatorrhea: Increased
Oxalate Solubility and Increased Colonic Permeability
- Mechanisms of Diarrhea and Steatorrhea in Patients with Bile Salt Malabsorption
- Diarrhea Caused by Fat Malabsorption
- Unabsorbed Nutrients Are Metabolized by Enteric Flora
- Fat Balance is the Test for Steatorrhea
- Detection of Steatorrhea
- Detection of Bile Salt Malabsorption
- Treatment of Fat Malabsorption
- Diagnosis of Major Causes of Fat Malabsorption
- Low-Fat Diet Decreases Diarrhea in Patients eith Fat Malabsorption
- Characteristics of Bile Salt and Fatty Acid Diarrhea Afet Ileal Resection
|